IMHO, I think you had the highlights in the right place, just needed more contrast. On the top, definitely more bright color. The sun is coming from the top just behind the animals, so a glow about the top of the mane would be good, moreso than I have here. I started out using the lasso tool and the render lights and ended up selecting larger areas, using the curve tool to lighten it and blending. The lower end is a bit strong on the shadow. So this is just a feeble attempt, but its what I'm thinking about. Your mane is wonderful. I just thought it should be lighter and more contrasty. I didn't lighten the center portion and perhaps it should have been slightly brighter.