I read somewhere that the wing of a bird is much like our arm - shoulder, elbow, wrist and fingers - The site you found is very interesting. I also found one that detailed how a designer was using his program and complex math to assign movement to the outer feathers depending on how it bent - way too much for me. I played around with morphs last night some - not a bad way to go since the bones deform the feathers - but one could take the feathers out of the bond influence, but then they wouldn't move with the wing tip.

Never Give Up!