The only differences I can see in the CR2 between bone_1 and bone_2  that seem to relate to this are highlighted in yellow on the comparison screenshots below - bone_1 on the left, bone_2 on the right.

Firstly, the twistY, jointX and jointZ are in a different position and order in the channels section and are named in a different manner

 

Secondly, the twistY, jointX and jointZ of bone_1 all have two extra parameters - flipped and primaryParm.

I've checked Kuroyume's unofficial CR2 spec but 2004 was a long time ago and it doesn't really give me any clues, except that the order of jointX and jointZ is important

While I wait for somebody who knows what they're talking about I may just try editing a copy of the CR2 to make the bone_1 joint parameters similar to the bone_2 ones... what's the worst that can happen ? ;o)

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The 3Dcheapskate* occasionally posts sensible stuff. Usually by accident.
And it usually uses Poser 11, with units set to inches. Except when it's using Poser 6 or PP2014, or when its units are set to PNU.

*also available in ShareCG, DAZ, and HiveWire3D flavours (the DeviantArt and CGBytes flavour have been discontinued).